Academic Appointments
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Professor, Biochemistry
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Member, Bio-X
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Faculty Fellow, Stanford ChEM-H
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Member, Wu Tsai Neurosciences Institute
Administrative Appointments
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Lead Investigator, Infectious Disease Initiative, Chan Zuckerberg Biohub (2017 - Present)
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Institute Scholar, Stanford ChEM-H (2014 - Present)
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Virginia & D.K. Ludwig Professor of Biochemistry, Stanford University School of Medicine (2014 - Present)
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President, Merck Research Laboratories, Merck & Co., Inc. (2003 - 2013)
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Executive Vice President, Merck Research Laboratories, Merck & Co., Inc. (2001 - 2002)
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Associate Head, Department of Biology, MIT (1999 - 2001)
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Investigator, Howard Hughes Medical Institute (1997 - 2001)
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Professor of Biology, MIT (1995 - 2001)
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Associate Investigator, Howard Hughes Medical Institute (1993 - 1997)
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Member, Whitehead Institute for Biomedical Research (1992 - 2001)
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Associate Professor of Biology, MIT (1992 - 1995)
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Assistant Investigator, Howard Hughes Medical Institute (1990 - 1993)
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Assistant Professor of Biology, MIT (1988 - 1992)
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Associate Member, Whitehead Institute for Biomedical Research (1988 - 1992)
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Whitehead Fellow, Whitehead Institute for Biomedical Research (1985 - 1988)
Honors & Awards
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Arthur Kornberg and Paul Berg Lifetime Achievement Award in Biomedical Sciences, Stanford University (2018)
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Member, National Academy of Sciences (1997)
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Member, National Academy of Medicine (formerly, Institute of Medicine) (2000)
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Member, National Academy of Engineering (2016)
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Fellow, American Academy of Arts and Sciences (2008)
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Doctor of Science, Honoris Causa, Pohang University of Science and Technology (2011)
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Presidents' Circle, The National Academies (2006)
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Harvey Lecture, The Harvey Society (2002)
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Fellow, Biophysical Society (1999)
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Fellow, American Association for the Advancement of Science (1999)
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Hans Neurath Award, The Protein Society (1999)
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Ho-Am Prize for Basic Science, The Samsung Foundation (1998)
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Fellow, American Academy of Microbiology (1997)
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DuPont Merck Young Investigator Award, The Protein Society (1994)
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Eli Lilly Award in Biological Chemistry, American Chemical Society (1994)
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NAS Award in Molecular Biology, National Academy of Sciences (1993)
Boards, Advisory Committees, Professional Organizations
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Medical Advisory Board, Howard Hughes Medical Institute (2016 - Present)
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Council, National Academy of Sciences (2015 - 2018)
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Scientific Advisory Board, Vaccine Research Center, NIAID, NIH (2014 - Present)
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MIT Corporation Visiting Committee, Department of Biology, MIT (2004 - Present)
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Board of Scientific Advisors, Jane Coffin Childs Memorial Fund (2015 - 2018)
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Advisory Council, Department of Molecular Biology, Princeton University (2015 - Present)
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External Scientific Advisory Board, Harvard Program in Therapeutic Science, HMS (2014 - Present)
Professional Education
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A.B., Cornell University, Chemistry (1979)
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Ph.D., Stanford University School of Medicine, Biochemistry (1985)