Molecular Genetic Pathology Fellowship


The Molecular Genetic Fellowship in the Department of Pathology currently has three funded fellowship positions per year.

This is an ACGME-accredited fellowship.

The Stanford Molecular Genetic Pathology program serves the adult and pediatric populations at Stanford and also sees referrals from Northern California and the rest of the nation.

The program is interdisciplinary and includes participation of faculty in the Departments of Pathology, Medicine, Pediatrics (Division of Medical Genetics) and Genetics. Laboratory rotations include formal training in Molecular Genetic Pathology, Biochemical Genetics, Cytogenetics, and Molecular Microbiology/Virology.

Fellows will be trained in assay development, quality assurance, and results interpretation in the Molecular Pathology Laboratory at Stanford and the Kaiser-Permanente Regional Molecular Diagnosis Laboratory, a large reference laboratory for the Kaiser system which offers a testing menu that is complimentary to that of Stanford.

Fellows are expected to complete a translational research project resulting in a clinical test for patient care during the fellowship.

Stanford University offers an outstanding environment for research opportunities in medicine, technology, and computational data sciences relevant to molecular pathology.

Eligible candidates for the Molecular Genetic Pathology fellowship must be board certified or board-eligible by the American Board of Pathology or the American Board of Medical Genetics.


Fellowship positions for 2021-2022 and 2022-2023 have been filled.

Applications will be reviewed on a rolling basis.


Please download, fill out and send the application along with the following: a CV, a personal statement, and three letters of recommendation. These should be submitted electronically to:

Mirela Puljic


James Zehnder, MD

Fellowship Program Coordinator

Gabby Barela

Current Fellows

Jack Tung, MD, PhD (2020-2021)
Emory University School of Medicine 

Julianna Weiel, MD (2020-2021)
University of Arizona College of Medicine 

Simon Chen, MD (2020-2021)
Washington University School of Medicine

Incoming Fellows

Maggie Black, MD (2021-2022)
Albert Einstein College of Medicine 

Jyoti Kumar, MD (2021-2022)
Ross University School of Medicine 

Lauren Lawrence, MD (2021-2022)
University of California Keck School of Medicine

Molecular Genetics Faculty

Helio Costa, PhD

Instructor in Pathology (Genomics & Molecular Genetic Pathology),
and Medicine (Data Management Systems)

Helio Costa, PhD, is a geneticist with expertise in genomics, molecular biology, molecular oncology, and bioinformatics. He is currently an Instructor within the Departments of Pathology and Biomedical Data Science at Stanford Medical School. Dr. Costa's research utilizes next-generation sequencing to develop new clinical genome and transcriptome profiling methods with the end goal of translating these tools to clinical diagnostic tests for implementation at Stanford Health Care. His research group is also interested in developing data science and machine learning methods to model and predict clinical outcomes and aid in clinical decision support. He is the founding director of the Stanford Clinical Data Science Fellowship where post-doctoral fellows engage in interdisciplinary clinical research and embed in health care workflows learning, building and deploying real-world health data solutions in the Stanford Health Care system. Additionally, he is an Attending Geneticist, and Assistant Lab Director of the Molecular Genetic Pathology Laboratory for Stanford Health Care. Dr. Costa received his BS in Genetics from University of California, Davis, his PhD in Genetics from Stanford University School of Medicine, and his ABMGG Clinical Molecular Genetics and Genomics fellowship training from Stanford University School of Medicine.

More information about Dr. Costa >>